reduce randomly selects species for inclusion in a reduced-richness
landscape based on the original. shuffle randomizes species locations
among sites.blender.reduce(landscape, richness)
blender.shuffle(landscape)data.frame for species (rows)
at sites(columns)data.frame based on landscapereduce without replacement. Species
are randomized by shuffle in order to preserve their occupancy
rates (row sums) but not local richness (column sums).