"Bio"The component of the operating model that controls biological dynamics, i.e., natural production.
NameCharacter. Identifying name
maxageInteger. The maximum age of the population age structure.
n_gInteger. Number of life history groups within a cohort. Life history groups (LHGs) are sub-units of a cohort that have different biological parameters, e.g., survival, but the egg production and smolt production in the next generation is calculated from the sum across life history groups. Default is 1.
p_LHGThe proportion of the total egg production assigned to each life history group within a cohort. For example,
if Bio@n_g <- 2, then Bio@p_LHG <- c(0.9, 0.1), then 90 percent of the egg production in the first population is assigned to the first life history group and
ten percent to the second LHG. Default is rep(1/Bio@n_g, Bio@n_g)
p_matureEither vector by age (length maxage) or an array with dimension [nsim, maxage, nyears+proyears]. The proportion mature by age.
SRrelCharacter, stock-recruit relationship for density-dependent smolt production from fry. Either "BH" (Beverton-Holt) or "Ricker".
Not used if habitat component is used. See Habitat object.
capacityVector length nsim. The asymptote of the Beverton-Holt stock-recruit function, or the Ricker maximum for
density-dependent natural smolt production from egg production. Units of smolts.
Not used if habitat component is used.
kappaVector length nsim. The adult productivity ratio for the stock-recruit function. Units of recruits per spawner.
Natural per-capita production of recruits as the population approaches zero (density-independent component).
In stage-based models, equivalent to the product of smolt productivity (smolts per spawner) and marine survival.
Not used if habitat component is used.
SmaxVector length nsim. The egg production that maximizes smolt production in the Ricker stock-recruit function. Units of eggs.
Equivalent to units of spawners if fec = 1 for all spawners. Not used if habitat component is used.
phiOptional parameter, vector length nsim. Unfished egg production per smolt. Units of egg per smolt.
The alpha parameter of the stock-recruit function will be the ratio of kappa and phi. In stage-based models,
this is the product of marine survival, fecundity, and proportion female. If not provided, phi will be calculated from Mjuv_NOS, p_mature, s_enroute, p_female, fec, and p_LHG corresponding
to the first year and weighted by life history groups. Not used if habitat component is used.
Mjuv_NOSEither vector by age (length maxage) or an array with dimension [nsim, maxage, nyears+proyears, n_g].
Natural mortality of immature natural origin fish.
To replicate the SAR parameter of a stage-specific model, set Mjuv_NOS[a] = -log(SAR) where a is the age class prior to maturation (and zero for all other ages).
fecVector by age (length maxage) or an array with dimension [nsim, maxage, nyears+proyears]. Female fecundity of natural origin spawners.
p_femaleNumeric. The proportion of females in the spawning population. Default is 0.5.
s_enrouteNumeric. Survival of escapement to the spawning grounds (for spawning and for broodtake). Default is 1.
Objects can be created by calls of the form new("Bio")
Various parameters can be stochastic (length nsim) or input as a single numeric
(value identical across all simulations).