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After calculating all pairwise distances between all tips, this function takes the variance across these values.
var_pair_dist(phy)
Variance in pairwise distance
phylo object or ltable
Webb, C., D. Ackerly, M. McPeek, and M. Donoghue. 2002. Phylogenies and community ecology. Annual Review of Ecology and Systematics 33:475-505.