sMLH is defined as the total number of heterozygous loci in an individual divided
by the sum of average observed heterozygosities in the population over the
subset of loci successfully typed in the focal individual.
Usage
sMLH(genotypes)
Arguments
genotypes
data.frame with individuals in rows and loci in columns,
containing genotypes coded as 0 (homozygote), 1 (heterozygote) and NA (missing)
Value
Vector of individual standardized multilocus heterozygosities
References
Coltman, D. W., Pilkington, J. G., Smith, J. A., & Pemberton, J. M. (1999).
Parasite-mediated selection against inbred Soay sheep in a free-living,
island population. Evolution, 1259-1267.